Halfords has reopened its branch in Broadbridge Heath, along with stores in Bexhill, Bognor Regis and Chichester.
The stores will have extensive social distancing measures in place, using an operating model that has been called Retail Lite, according to the firm.
A spokesman added: “Having been granted essential retailer status, 335 of Halfords’ 446 retail stores have until now been operating under a Dark Store model during the lockdown.
“This means that customers have been unable to browse and instead have been placing orders with Halfords colleagues outside the front of the store.”
Customer numbers in the store will be limited with safety notices and floor markings to remind customers about social distancing.
There will be a cleaning station at the front of the store, with supplies of hand sanitiser and wipes alongside other measures to keep staff and customers safe.
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