Environmental Health is the latest public service to be shared by Rother and Wealden Partnership as the twocouncils continue to cut costs.
The joint council initiative is set to save £250,000 a year through the agreement.
Administrative support for the shared service will now be based at Bexhill in Rother District.
Residents can still continue to contact their own council directly for services including alcohol and taxi licencing, noise and smoke pollution, health and safety enforcement, food safety inspections and environmental monitoring.
Cllr Bob Standley, leader of Wealden District Council, said: “Staff have worked hard to ensure there is a seamless transition to this new shared service which further strengthens the close working between Rother and Wealden for the benefit of our residents
Rother District Council leader Cllr Carl Maynard said: “The creation of a shared service for Environmental Health is our biggest step forward yet.
Wealden customers should contact 01323 443322 or visit www.wealden.gov.uk